I wonder if this model just needs time after boot up to recognize the HDD. I have the 670 and sometimes I had to do a hard reset since it still showed couple of connections that were turned off. Try to reinstall old firmware from website.
DO THIS: Go to Settings, Others and select Initialize, Factory Set. Turn off unit, unplug unit for a minute, plug in, turn back on. Configure your network settings, download 404E update again, repeat above procedure after update turns off player.